they have a singular knee extended into a fork-like projection,/, which in the flower lies just below the bright yellow anthers of the short stamens. The lower portion of the long stamens takes a backward curve from the fork carrying the pores of its anthers far from those of the short stamens. This arrangement is most beautifully adapted, as was first pointed out by Fritz Ji a.t
MAOEller, for the cross-fertilisation flower (diagrammatic)of melastoma.
a , , IN SECTION.a p, PETAL ,* a1, ANTHEU
01 the plant. of SHORT eTAMENS ; AE , FORK OF 1 OXG
The bees invariably made stamens ; a\ anther of long sta-
n 1*1 * i A tENS ; C. FILASI ENT OF LONG ANTHEU ;
lor the bright yellow platiorm ht, stigma of pistil ; A , ovary. offered by the bunch of short
stamens (perhaps because they do not perceive from a distance the pink pistil and long stamens projected against the pink corolla), and invariably received the pistil between their legs, their feet settling also on the adjoining fork of the long stamens. The instant effect of this is to collect the whole of the long stamens into a bunch, and to depress their anthers downwards and away from the body of the bee, while the stigma of the pistil (which hangs down close to the pores of the long-stamened anthers) remains in constant contact with its ventral side. At the moment of the beea s departure, the hooks on its feet, by pulling on the fork of the long stamens, raise their anthers, bringinga now that there is no fear of producing self-fertilisation of the planta their tips in a collected bunch into contact with its sides and abdomen. Long after I had made these observations, w hile working in the laboratory of the Buitenzorg Gardens, I)r. Burek pointed out to 1X16 a fact of considerable importance which I was able to verify for myself, that there was in very closely allied species of this amily a great difference in the shape of the pollen of the two orms of anther ; that while pollen of both shapes was found on the pistil, that from the long stamens alone seemed fertile. W e Could not detect any pollen tubes (which are emitted when the Pollen is fecundating the plant) emanating from the pollen of t e short-stamened anthers.
The reason why some organ of a plant or animal has assumed, as it were, an abnormal form, is not always easy to discover ;