A NATURALIST'S WANDERINGS
its anther erect on the posterior part of the column. Underneath is the stigma, of a roughly square shape, its upper rim standing erect in front of the pollinia, rising to about one third of their height as a triangular eminence, which corresponds with the front margin of the rostellar platform. It is not in every flower that the shape of the stigma can be seen well, for the stage presently to be described begins very soon, often before the flower is expanded ; and only by the examination of a very large series have I been able to follow the modifications that have occurred.
Concurrent with or even before the commencement of the rotation of the anther into its normal position some in-
no. 20.a ARUNDINA SPECIOSA, SHOWING FIG. 21.a ARUNDINA SPECIOSA, BUD SHOW-
G, STYLARY CANAL.
fluencea which I do not knowa causes the upper margin of the stigma to become inverted close down the posterior wall of the stylary canal, as seen in Fig. 17, and in longitudinal section opened from behind in Fig. 20, where the rostellum is seen hanging down the canal as a narrow band. Fig. 21 represents a very young bud, in which, though the pollinia had scarcely begun to rotate, the stigma had become already much modified, and is in waiting for the rotation of the pollinia. Along with this invagination of the upper margin of the stigma (the rostellum) its lower lip is in consequence dragged (?) upwards. Dissections of the column showed that the rostellum goes on elongating down the stylary canal, as in Fig. 20, while the pollinia, slowly continuing to rotate downwards, finally precipitate themselves into the stigma, whose flap-like margins embrace the
A SECTION OF COLUMN OPENED FKOM REHIND ; C, TOP OF UPPER MARGIN OF STIGMA (CORRESPONDING TO
rcstellum); c2, portion of stigma;
ING THE LPPER RIM OF STIGMA ALREADY INVERTED DOWN THE STYLAHY CANAL; LETTERS AS IN FIG. 17.