narrow channel by which the viscid matter of the stigma reaches the anther. In examining an advanced bud, the viscid matter of the stigma is seen to be in large quantity and rather liquid. It increases with the growth of the flower till it overflows,a often before the bud opensa and, immediately on its opening, inundates the pollinia, which now increase in size, and either avalanche downwards, sometimes quite obliterating the rostellum (Fig. 2, p. 86); or, while retaining their position in the anther, emit their tubes over the narrower portion of the rostellum into the stylary canal. Very often both anther and stigma become quite filled up by the multitude of pollen-tubes and by the swollen pollinia. All these plants produced large and well-filled seed-capsules on every flower ; but I
PIG. 5.a PHAJUS BLUMET, SHOWING THE FIG. 6.a 1HAJUS BLUMEI, SHOWING A
never saw an insect visit the plants during all my observations, although the plants were situated where I could inspect them constantly throughout the day or night.
Of flowers of the second form, I examined many more examples. Here there is no rostellum, nevertheless the boundaries of the stigma are quite distinct (Figs. 3, 4, p. 86). On examining a young bud, the anther (enclosing the pollinia) is seen standing vertically erect on the top of the columna i.e. of the detached column, without reference to its position in the flowera forming as it were a pointed extension of it, and attached to it by its niinute filament. As the flower progresses in growth, the anther-cap ruptures and rotates forward. When it has descended through about 90A , it occupies (Fig. 5) the position which, if it possessed a rostellum, it would naturally retain : but, having
ANTHER ROTATED 1 OWN WARDS ; A, C, AS IN FIG. 3.
MORE ADVANCED STAGE THAN FIG. 5 ; THE ANTHER-CAP A, HAS OPENED ; B, SWOLLEN POLLINIA ; C, E, AS IN FIG. 2 ; K, TIP OF ANTHEU-CAP.