can effect it) ; that the pollen grains exsert their pollen tubes while still in the anthers ; and that both the external and the internal surfaces of the lobes of the pistil are oovered with papillae, indicating that these surfaces are functionally active.
I have never observed these flowers approached by the ants that infest the interior, nor by any other insect, which to gain admission to the flower, even if open, must be very small indeed. The anthers and the pistil do not seem to reach maturity together, yet it would seem that self-fertilisation alone can take place ; perhaps the tubes of the pollen grains which fall to the bottom of the corolla manage to reach the lower lobes of the pistil and produce fecundation.
The seeds I planted germinated with great freedom, and I cultivated quite a number of young Myrmecodia, whose growth I watched with the greatest interest. Many of them I kept quite isolated from the interference not only of the Pheidole javana, which seems to be the only species of ant which lives in these plants in their native state, but of all other species, and I was surprised to find that from their very earliest appearance this curious galleried structure arose without the presence of the ants, and that the plants continued to grow and thrive vigorously in their absence as long as I cultivated them. Some bulbs had a single canal reaching to their centre from a round
orifice opening generally close to the little tap-root ; others presented one or two loculi in the interior, without any communication at first with the exterior, partially full of a spongy substance looking like its own degenerated tissue. These chambers invariably developed a spongy pitha which in a section it was not difficult to trace out in advance in the still fleshy substancea towards and to open at
10csj. n4.____, ,1 , a YOUNG MYRMECODIA, AND
last at one or more spots on the exterior ot section of a somewhat the bulb. Secondary galleries, arising in older one. the same manner as the primary, soon formed communicating channels, extending with age, throughout the whole of the growing bulb. At a later period, in Amboina, where the Myrmecodia and the Hydnophytum were very abundant, I found many specimens containing a large central and quite isolated chatpber full of watera not rain-watera round which